The species richness of a community is the number of species in it. Richness alone ignores abundance: a wood with a hundred oaks and one of each of nine other trees is less diverse than one with ten of each. The Shannon index
where is the fraction of individuals belonging to species , rises with both richness and evenness; it is when all species are equally abundant and near zero when one dominates. The evenness is . Diversity generally rises from the poles to the tropics, with area, with time since the last disturbance, and with the productivity of the site up to a point.
Examples
Example 26.14 (Two woods)
Wood A: 100 oaks and one each of nine other trees; , evenness . Wood B: ten each of the same ten species; , evenness 1. Same richness, five times the diversity.